In RPGs up until Moon: Remix RPG Adventure, the “hero” has been a hero. However, the “Hero” who appears in the world of Moon kills innocent monsters (referred to as “animals” in the game), breaks into other people’s homes to burgle various items and harass residents, and is generally troublesome whenever he appears. The role of the protagonist, “Boy,” is to free the souls of the animals that the hero has slain and touch the hearts of the world’s strange, warmhearted inhabitants. To grow, one must level up one’s Love. The major theme is woven into the story, so one must be thoughtful to see the game’s true ending.

Moon begins with the protagonist, a small boy, playing a new RPG called "Moon" (a.k.a. "Fake Moon") on his "Gamestation." The game begins with the player controlling the Hero of Fake Moon in a 10-minute game-within-a-game, Fake Moon being something of a parody of Japanese RPGs of the 16-bit era.[7] Convoluted JRPG stories are skewered by the minutes of nonsensical backstory, which Boy skips through before the player can read it. Queen Aphrodite has been abducted and taken to the moon. The perpetrator, Dragon, will wreak millions of calamitous years upon the people of Love-De-Gard with her power. Yet, the people have produced a hero who must travel to Dragon Castle and destroy the beast. After playing through a few typical RPG scenes (random battles, an airship sequence, etc.), the boy is ordered by his mother to go to bed and obediently does so. However, the television on which he was just playing Fake Moon switches back on by itself, and the boy is sucked into the world of Moon, a land called "Love-de-Gard." Its people and its story resembles Fake Moon’s.